Arbutoideae

Arbutoideae
	Arbutus menziesii
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Ericales
Family:	Ericaceae
Subfamily:	Arbutoideae; Nied.
Type genus
	Arbutus;
Genera
	Arbutus; Arctostaphylos; Arctous; Comarostaphylis; Ornithostaphylos; Xylococcus;

The Arbutoideae are a subfamily in the plant family Ericaceae. Phylogenetic analysis supported all genera of the subfamily as monophyletic, except Arbutus.^[1] Moreover, it was suggested that the non-sister relationship between Mediterranean and North American species may be explained by a once widespread distribution in the Northern hemisphere before the Neogene.^[1]

The genera Arbutus, Arctostaphylos, Comarostaphylis form a particular type of mycorrhizal symbiosis with the fungus, Arbutoid mycorrhiza, which resembles ectomycorrhizas.^[2]^[3]

Genera List[edit]

Image

Genus

Living species

Arbutus L.

Species	Common name	Range
andrachne	Greek strawberry tree
canariensis	Canary madrone
pavarii	.
unedo	Strawberry tree
arizonica	Arizona madrone
bicolor	.
madrensis	.
menziesii	Pacific madrone
mollis	.
occidentalis	.
tessellata	.
xalapensis	Texas madrone

Arctostaphylos Adans.

Subgenus Micrococcus
- Sect. Micrococcus

Species	Common name	Range
mendocinoensis	Pygmy manzanita
myrtifolia	Ione manzanita	California (Amador, Calaveras Counties)
nissenana	Nissenan manzanita	California (coastal and inland ranges north of San Francisco Bay)
nummularia	Glossyleaf manzanita	California (Mendocino County)

Subgenus Arctostaphylos, which has 3 sections:
- Sect. Arctostaphylos

Species	Common name	Range
alpina	Alpine bearberry
bakeri	Baker's manzanita	California (Sonoma County)
densiflora	Sonoma manzanita	California (Sonoma County)
edmundsii	Little Sur manzanita	California (Monterey County)
franciscana	Franciscan manzanita	California (San Francisco County)
gabrielensis	San Gabriel manzanita	California (Los Angeles County)
glauca	Bigberry manzanita	California and Baja California
hispidula	Gasquet manzanita	Coastal mountain ranges of southern Oregon and northern California
hookeri	Hooker's manzanita	California
insularis	Island manzanita	California (Santa Cruz Island)
klamathensis	Klamath manzanita	California (Klamath Mountains)
manzanita	Common manzanita, whiteleaf manzanita	California (Coast Ranges and Sierra Nevada foothills)
mewukka	Indian manzanita	California (Sierra Nevada)
nevadensis	Pinemat manzanita	California
parryana	Parry manzanita	California (southern)
patula	Greenleaf manzanita	Western North America
pumila	Sandmat manzanita	California (Monterey County)
pungens	Pointleaf manzanita	Southwestern United States and to northern and central Mexico
rudis	Shagbark manzanita	California (southern central coast)
stanfordiana	Stanford's manzanita	California (Outer North Coast Ranges north of the San Francisco Bay Area)
uva-ursi	Bearberry	Europe, Asia, North America
viscida	Sticky manzanita, whiteleaf manzanita	California and Oregon

- Sect. Foliobracteata

Species	Common name	Range
andersonii	Santa Cruz manzanita
auriculata	Mount Diablo manzanita
canescens	Hoary manzanita
catalinae	Santa Catalina Island manzanita
columbiana	Hairy manzanita
confertiflora	Santa Rosa Island manzanita
cruzensis	La Cruz manzanita
glandulosa	Eastwood manzanita
glutinosa	Schreiber's manzanita
hooveri	Hoover's manzanita
imbricata	San Bruno Mountain manzanita
luciana	Santa Lucia manzanita
malloryi	Mallory's manzanita
montaraensis	Montara manzanita
montereyensis	Monterey manzanita
morroensis	Morro manzanita
nortensis	Del Norte manzanita
obispoensis	Serpentine manzanita
osoensis	Oso manzanita
otayensis	Otay manzanita
pajaroensis	Pajaro manzanita
pallida	Pallid manzanita
pechoensis	Pecho manzanita
pilosula	La Panzo manzanita
purissima	La Purissima manzanita
refugioensis	Refugio manzanita
regismontana	Kings Mountain manzanita
silvicola	Bonny Doon manzanita
tomentosa	Woolyleaf manzanita
virgata	Bolinas manzanita
viridissima	Whitehair manzanita
wellsii	Wells' manzanita

- Sect. Pictobracteata

Species	Common name	Range
pringlei	Pringle manzanita

Unassigned

Species	Common name	Range
rainbowensis	Rainbow manzanita
gabilanensis	Gabilan manzanita
ohloneana	Ohlone manzanita

Arctous (A.Gray) Nied.

Species	Common name	Range
alpina (L.) Nied.
microphyllus C.Y.Wu
ruber (Rehder & E.H.Wilson) Nakai

Comarostaphylis Zucc.

Species	Common name	Range
arbutoides
discolor
diversifolia
glaucescens
lanata
longifolia
mucronata
polifolia
sharpii
spinulosa

Ornithostaphylos Small

Species	Common name	Range
oppositifolia	Baja birdbush

Xylococcus Nutt.

Species	Common name	Range
bicolor	Mission manzanita

References[edit]

^ ^a ^b Hileman LC, Vasey MC, Parker VT (2001). "Phylogeny and biogeography of the Arbutoideae (Ericaceae): implications for the Madrean–Tethyan hypothesis". Systematic Botany. 26 (1): 131–143. doi:10.1043/0363-6445-26.1.131 (inactive 31 January 2024).{{cite journal}}: CS1 maint: DOI inactive as of January 2024 (link)
^ Osmundsen TW, Halling RE, den Bakker HC (2007). "Morphological and molecular evidence supporting an arbutoid mycorrhizal relationship in the Costa Rican páramo". Fungal Diversity. 17 (3): 217–222. Bibcode:2007Mycor..17..217O. doi:10.1007/s00572-006-0098-x. PMID 17216498. S2CID 19084561.
^ Smith SE, Read D (2008). Mycorrhizal Symbiosis (3rd ed.). Amsterdam; Boston: Academic Press. ISBN 978-0-12-370526-6.

External links[edit]

Data related to Arbutoideae at Wikispecies
Arbutoideae in the Flora of North America

This Ericaceae article is a stub. You can help Wikipedia by expanding it.

[Hileman2007-1] Hileman LC, Vasey MC, Parker VT (2001). "Phylogeny and biogeography of the Arbutoideae (Ericaceae): implications for the Madrean–Tethyan hypothesis". Systematic Botany. 26 (1): 131–143. doi:10.1043/0363-6445-26.1.131 (inactive 31 January 2024).{{cite journal}}: CS1 maint: DOI inactive as of January 2024 (link)

[Osmundsen2007-2] Osmundsen TW, Halling RE, den Bakker HC (2007). "Morphological and molecular evidence supporting an arbutoid mycorrhizal relationship in the Costa Rican páramo". Fungal Diversity. 17 (3): 217–222. Bibcode:2007Mycor..17..217O. doi:10.1007/s00572-006-0098-x. PMID 17216498. S2CID 19084561.

[SmithRead2008-3] Smith SE, Read D (2008). Mycorrhizal Symbiosis (3rd ed.). Amsterdam; Boston: Academic Press. ISBN 978-0-12-370526-6.

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